Papers in Palaeontology

Rostral and body shape analyses reveal cryptic diversity of Late Jurassic batomorphs (Chondrichthyes, Elasmobranchii) from Europe
Türtscher J, Jambura PL, Villalobos-Segura E, López-Romero FA, Underwood CJ, Thies D, Lauer B, Lauer R and Kriwet J
The fossil record of chondrichthyans (chimaeras, sharks, rays and skates) consists largely of isolated teeth, with holomorphic specimens being extraordinary exceptions. However, numerous of these more or less completely preserved specimens are known from several Upper Jurassic deposits of Europe, enabling detailed analysis of their morphology. Batomorphs (rays and skates) resembling modern guitarfishes and wedgefishes (Rhinopristiformes) are among the most common Jurassic chondrichthyans found, but they have been only sporadically studied up to now, resulting in large knowledge gaps concerning their taxonomy and phylogeny. Here, we present the most detailed revision of Late Jurassic holomorphic batomorphs to date, quantitatively analysing body proportions of specimens from Germany (Solnhofen Archipelago), France (Cerin) and the UK (Kimmeridge), using both geometric and traditional morphometrics. Furthermore, we identify qualitative morphological characters for species discrimination, to clarify the taxonomic identity and diversity of Late Jurassic batomorphs based on holomorphic specimens. Our results support the validity of and , as well as that of the previously doubtful . Moreover, we describe , a new taxon, which has hitherto been considered to be a large-sized morphotype of . Our results highlight that the diversity of holomorphic batomorphs during the Late Jurassic was greater than previously thought, and suggest that this group was already well-established and diverse by this time. This study thus provides vital information about the evolutionary history of Late Jurassic batomorphs and has direct implications for batomorph species that are based on isolated teeth only.
An Eocene paraclupeid fish (Teleostei, Ellimmichthyiformes) from Bolca, Italy: the youngest marine record of double-armoured herrings
Marramà G, Bannikov AF, Kriwet J and Carnevale G
A new double-armoured herring of the clupeomorph order Ellimmichthyiformes, gen. et sp. nov., is described herein based on a single partially complete specimen from the early Eocene Pesciara site of the Bolca Konservat-Lagerstätte, north-eastern Italy. The fossil documented herein exhibits a unique combination of features (e.g. ornamentation of the skull bones; medial fusion of the contralateral halves of the neural arches of abdominal vertebrae; teeth on endopterygoid, parhypural fused to the first preural centrum; presence of a short series of 6 (or 7) predorsal scutes increasing in size posteriorly; postpelvic scutes bearing very prominent spines), which clearly supports its recognition as a new genus and species of the family Paraclupeidae. Evidence is provided to show that gen. nov. is most closely related phylogenetically to the Early Cretaceous genus . The new taxon described herein represents the youngest marine occurrence of the family Paraclupeidae and, more generally, of the order Ellimmichthyiformes, suggesting that the shallow marine settings of the Tethys might have favoured the persistence of certain fish lineages that were severely affected by the end-Cretaceous mass extinction.
Anatomy of the Ediacaran rangeomorph
Dunn FS, Wilby PR, Kenchington CG, Grazhdankin DV, Donoghue PCJ and Liu AG
The Ediacaran macrofossil Ford is perhaps the most iconic member of the Rangeomorpha: a group of seemingly sessile, frondose organisms that dominates late Ediacaran benthic, deep-marine fossil assemblages. Despite exhibiting broad palaeogeographical and stratigraphical ranges, there have been few morphological studies that consider the variation observed among populations of specimens derived from multiple global localities. We present an analysis of that evaluates specimens from the UK, Canada and Russia, representing the largest morphological study of this taxon to date. We describe substantial morphological variation within and present a new morphological model for this species that has significant implications both for interpretation of rangeomorph architecture, and potentially for existing taxonomic schemes. Previous reconstructions of include assumptions regarding the presence of structures seen in other rangeomorphs (e.g. an internal stalk) and of homogeneity in higher order branch morphology; observations that are not borne out by our investigations. We describe variation in the morphology of third and fourth order branches, as well as variation in gross structure near the base of the frond. The diagnosis of is emended to take account of these new features. These findings highlight the need for large-scale analyses of rangeomorph morphology in order to better understand the biology of this long-enigmatic group.
AN EARLY MIOCENE DEEP-WATER DECAPOD CRUSTACEAN FAUNULE FROM THE SLOVENIAN PART OF THE STYRIAN BASIN, AND ITS PALAEOENVIRONMENTAL AND PALAEOBIOGEOGRAPHICAL SIGNIFICANCE
Gašparič R and Hyžný M
A new decapod crustacean faunule is described from the early Miocene of the Slovenian part of the Styrian Basin. The Ivnik Beds exposed at the Činžat locality contain seven species: (Axiidea: Callianassidae), Lepidophthalmus paratethyensis sp. nov. (Axiidea: Callianassidae), (Gebiidea: Laomediidae), (Brachyura: Chasmocarcinidae), (Brachyura: Goneplacidae), sp. nov. (Brachyura: Mathildellidae) and sp. nov. (Brachyura: Retroplumidae). Numerous specimens of well-preserved permitted its redescription and re-assignment of its familial placement to Chasmocarcinidae. sp. nov. constitutes the first fossil occurrence of the genus known to date. The decapod association, as well as other faunal elements, suggests low-energy deep-water depositional environment with epibathyal water depth of more than 125 m. The studied locality is situated in the Ribnica-Selnica graben filled with sediments once deposited in the Central Paratethys sea. Based on the affinities of decapod genera of the Central Paratethys and the Proto-Mediterranean, we conclude that the exchange of decapod faunas between these regions was probably regulated by an anti-estuarine circulation permitting an easier incursion of species from the Proto-Mediterranean into the Paratethys and simultaneous hindering the Paratethyan endemics () from entering the Mediterranean.